Five girls came to a training school to become voice actresses holding each thought in her mind. Momoko - she wanted to be a voice actress to meet her favorite voice actor. Rinka - she is an armature voice actress, but she didn't have special purpose to become voice actress. Yurika - she likes anime very much. Tsubasa - she was a girl in love, and acted only as a boy. Amane - she found a deadlock as a actress and tried to find another way. They managed to get over the difficulty to become voice actress although they couldn't perform as main characters yet. There is a love romance with a boy works for an anime company, and you can catch a glimpse of situations of anime industries.
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After becoming a voice actress trainee, Momoko's new life begins.
Momoko finds living in Tokyo hard as she has no money. She tries to find a suitable part-time job.
Momoko and her friends help out in a stage debut, but unexpectedly, an accident appears.
Rinka, one of Momoko's friend, loses confidence in being a voice actress.
Tsubasa is a tomboy who wants a male role as a voice actress. During an event, Tsubasa learns what it feels like when a girl is going to a date in order to practice a female role. She goes on a date with her senpai who she has a crush on.
Yurika remembers her promise with Atari back when they were little kids, when Atari pinky-promised Yurika that when she became a voice actor, he'd draw the animations. When Yurika finds out that Atari has already had his debut in animating, she rushes to have her own debut, leading her to be tricked by a fake company and quitting Lambda 8. Momoko, Atari and Daimon find Yurika, in a bathing suit being harassed but saves her in the end. Momoko finds out Yurika might be in love with Atari.
Amane's father forbids her to continue on training to become a voice actor and takes her home. Momoko along with the rest of the girls and Kiyoka try helping Amane to escape, and in the end Amane's father realizes that voice acting is just as good as acting and he lets her continue on training.
Momoko is invited to practise for the Special Promotion with her idol Minato but because she knew that Yurika and Atari were going on a date on the same day, she becomes jealous and doesn't pay attention during her practice with Minato. Back at the group home, Runa tells Momoko that Momoko's in love with Atari even though Momoko won't be able to prove that to herself yet.
Amane was declined from her job and was asked to instead act as one of the characters of "Child Girls" in a promotional act. Momoko and the others were also asked to be Amane's supportive characters on stage. When things go wrong, Momoko uses her adlib skills to save the show!
Momoko and the others are given the opportunity to try out as the main character in an anime. They all practise how to say the lines and Momoko wins the part. But in the end, the company cancels because the leader of the company is a fan of Yuumi and wanted Yuumi to do the part at all cause.
While everyone else has a job, Momoko has none until she was asked to play as "Chiiko" in "Child Girls". Due to it being her first job, she was nervous for being the only rookie in the studio. She was corrected many times and was hopeless until Minato tells her that everyone started off as a rookie. Momoko realizes that she should be happy for the opportunity and keep trying so she took a lot of retakes until they were happy. Meanwhile, Atari also was given his debut - to draw for Child Girls!
Lambda 8 is going bankrupt, so to fix that, they assembled Momoko, Amane, Yurika, Tsubasa and Rinka to become the "Sister x Sisters"! Before the performance, things go wrong as people assumed that the Sister x Sisters were little sister characters, not Nuns, so Momoko and the team create a way to work it like usual. It pulls off, the show goes well and Lambda 8 stays in business.
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